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Quick highlights from the Buffalo Bills' exhibition loss to the Tampa Bay Buccaneers in Ralph Wilson Stadium ...

What it means: Oy to the vey. The Bills' starting offense looked atrocious, and their first-string defense was middling. The Bills' starters were outscored, 24-0, by the Bucs' starters at halftime. The Bills are 1-3 with one preseason game left.

Starters finally score: The Bills' first-team offense scored its first touchdown on its 19 preseason possession, then scored again on its 20th possession. Against the Bucs' backups, EJ Manuel connected with Mike Williams for a 14-yard touchdown on the first drive of the second half. Fred Jackson scored on an 8-yard touchdown run.

QB report: Manuel's fourth pass ended the opening drive with an interception. He also lost a fumble the Bucs returned for a touchdown. He was 19 of 28 for 198 yards and one touchdown with one interception. The Bucs sacked him four times. In the third quarter, Manuel was 10 of 10 for 131 yards against the Bucs' backups. ... Dennis Dixon was the surprise selection to quarterback the second unit and came in at the beginning of the fourth quarter. Dixon played one series. He dropped the ball three times while running, but he was ruled down by contact twice. Dixon's lone attempt was incomplete. ... Jeff Tuel finished the game. He completed eight of his 14 passes for 90 yards. He had an interception wiped out by roughing-the-passer penalty. Thad Lewis didn't play.

Play of the game: Bills gunner Duke Williams delivered a perfectly timed, skeleton-jolting hit on Bucs punt returner Solomon Patton with about seven minutes left in the second quarter.

Steven Means chips in for Bucs: The defensive end from Grover Cleveland High and the University at Buffalo played after missing practice time this week. Means chased Manuel out of bounds for a 7-yard sack in the third quarter.

Bills inactives: Wide receiver Sammy Watkins (ribs), tight end Tony Moeaki (hamstring), guard Chris Williams (back), defensive tackles Kyle Williams (leg) and Alan Branch (undisclosed), cornerbacks Bobby Felder (groin) and Mario Butler (ankle) and safety Jonathan Meeks (neck).

Preliminary injury report: Bills safety Da'Norris Searcy left the game with a stinger and did not return. Wide receiver Marquise Goodwin departed with a neck injury just before halftime. Tight end Lee Smith was knocked out with a foot injury.

Coming up: NFL teams must cut down their rosters to 75 players by 4 p.m. Tuesday. The Bills wrap up their preseason schedule Thursday night against the Detroit Lions.